Will Arnett
Lone Gunslinger Vulture
Peter Ackerman
Dung Beetle Dad
Caitlin Rose Anderson
Glypto Boy Billy / Beaver Girl
Connor Anderson
Rhino Boy / Beaver Boy
Nicole DeFelice
Start Girl
Debi Derryberry
Diatryma Mom
Jason Fricchione
Molehog Grandpa / Shovelmouth Male
James Godwin
Traffic Vulture
Clea Lewis
Female Mini Sloth / Dung Beetle Mom
Brian Scott McFadden
Glyptodon
Jansen Panettiere
Shovelmouth Boy
Gregory Romano
Start Boy 1
Manoela Scarpa Saldanha
Molehog Kid 1
Sofia Scarpa Saldanha
Molehog Kid 2
Reyna Shaskan
Diego's Bird Girl
Cindy Slattery
Aardvark Mom
Emily Anderson
Additional Voices
Jack Angel
Additional Voices
Shane Baumel
Additional Voices
Bob Bergen
Additional Voices
Claudia Besso
Additional Voices
Kyle J. Bogert
Additional Voices
Rodger Bumpass
Additional Voices
Paul Butcher
Additional Voices
Blanca Camacho
Additional Voices
John Cygan
Additional Voices
Jennifer Darling
Additional Voices
Olivia DeLaurentis
Additional Voices
Harrison Fahn
Additional Voices
Bill Farmer
Additional Voices
Crispin Freeman
Additional Voices
Spencer Lacey Ganus
Additional Voices
Jess Harnell
Additional Voices
Tim Kirkpatrick
Additional Voices
David H. Kramer
Additional Voices
Sherry Lynn
Additional Voices
Danny Mann
Additional Voices
Laura Marano
Additional Voices
Madeleine Martin
Additional Voices
Matt McCarthy
Additional Voices
Mickie McGowan
Additional Voices
Laraine Newman
Additional Voices
Jan Rabson
Additional Voices
Zack Shada
Additional Voices
Thomas Sharkey
Additional Voices
Ross Simanteris
Additional Voices
Rachel Diaz-Stand
Additional Voices
Ariel Winter
Additional Voices
Victoria Allegranza
Bird Girl